Papers written by QERC Students and visiting biologists are arranged here by topic. 

The majority of these papers are written in English, those with Spanish translations are noted (ENG/ ES). 

We are currently working to digitize the QERC library of papers and presentations. 

If you have an urgent request, or are the author of a paper you would like to see listed here please contact the Field Station Manager. 


  • Preliminary Report of the Oxford University Expedition to Costa Rica 1991
  • Lauracea Research Project, by Colin Berg 1991
  • Bromeliads in the Tropical Rainforest, by Wendy Huckaby 1993
  • Plant Species Diversity in the Valley of San Gerardo de Dota, by Sara VandenBosch 1994
  • Phenology of Lauraceae in San Gerardo de Dota...a Continuing Study, by Christine L. Duff 1994
  • A Base-Line Study of Extractivism Potentials at the QERC in the San Gerardo de Dota Valley in Costa Rica, by Angela E. Cheney 1995 
  • Light Intensity and Slope/ Aspect Correlations of the East and West Valley Walls of San Gerardo de Dota, by Kristen Borchers 1995
  • Medicinal Plants in the Valley of San Gerardo de Dota, by Beth A. Dieleman 1996
  • A study of the Number of Lauraceae Trees Located in the Rio Savegre Valley of Costa Rica, by Michelle Fedroza 1996
  • Percentages of Flowers and Fruits on Lauraceae Trees in the Cloud Forest of the Cordillera Talamanca, by Angelica M. Vasquez 1997
  • Medicinal Plants of the San Gerardo de Dota Cloud Forest Community, by Jennifer Jacobson 1997
  • Tree Fern Density and Distribution in San Gerardo de Dota, by Lisa Tebben 1997
  • A Study of the Effect of Light on the Number of Species in a Micro-ecosystem, by Nathan Daniel 1997
  • Phenological Profile of Lauraceae with Respect to Natural and Human Impact: a Comparative Analysis in San Gerardo de Dota, Costa Rica, by Colleen James 1998
  • Tree Ferns of San Gerardo de Dota: A Relationship Between Environmental Conditions and Distribution of dicksonia and sphaeropteris Species, by Jamie Pottier 1998
  • Heliconia in the Rio Savegre, by Connie Rush 1999
  • A Comparative Study - Secondary Growth in the Tropical Cloud Forest, by Sarah Allen 1999
  • Successional Study of Pasture Area in the San Gerardo de Dota Valley, Costa Rica, by Rebecca Rosaler 1999
  • Light Gap Analysis: Life at Ground Level, by Lisa Peterson 1999
  • Ethnobotany: Exploring the Potential of the Rio Savegre Valley, by Nicole Lang 1999
  • Successional Baseline Study of a Lower Montane Forest, by Ben Lassonde 1999
  • A Comparison of the Lauraceae Phenology, by Eva Jackson 1999
  • A Survey of Bacteria and their Antagonistic Properties in Bromeliads Experiencing Different Light Regimes in a Neotropical Montane Forest, by Melanie S. Spoor 2000
  • The Population of Heliconia in the QERC Stream Systems, by Ali Harris 2000
  • Ferns are a Sight for Sori (Sore Eyes), by Rebecca Florence Van Ee 2001
  • Two Tree Fern Populations and Their Soils, by Lesley Ediger 2001 
  • A Study of the Natural Climates of Presto acuminate and Genome hoffmanniana in a Cloud Forest at San Gerardo de Dota, by Lizabeth Larson 2002
  • A Comparative Study of Epiphyte Load on Trunks of Cloud Forest Oak Trees, by Joseph Regan Hutson 2002
  • Asteraceae: A Family of Flowers and Medicinal Uses in Costa Rica, by Candace M.R. Murray 2002
  • A Comparative Study of Natural and Unnatural Light Gaps in the San Gerardo de Dota, by Aaron Fouch 2002
  • Tree Fall Light Gaps: Percent Area and Average Size in the QERC Tropical Montane Rain Forest, by Brad Christoffersen 2002
  • Tropical Ecology and Mistletoe Survey, by Joanna Rice 2002
  • A Study of Wildflowers Growing Along the Banks of the Savegre River and their Antimicrobial Properties, by Kelly Pound 2003
  • Density and Diversity of Epiphytic Cover on Quercus Trees in a Tropical Montane Rainforest in Costa Rica, by Clark Wenger 2008
  • The Relationship Between Fruit Production and the Physical Environment of the Family Lauraceae  in San Gerardo de Dota, Costa Rica by Emily Ogle
  • Affects of Habitat Variation of Flower Production in Epiphytic Species of Ericaceae in the Rio Savegre Valley, Costa Rica, by Katherine Field 
  • Abiotic Factors Influencing the Growth of Tank Bromeliads in Upper Savegre Valley of Costa Rica, by Brittany Solis
  • Medicinal Plants of San Gerardo de Dota, by Holly Corwin
  • A study of Epiphytic Life on Quercus hosts in Primary and Secondary Forests Within the QERC Forest, San Gerardo de Dota by Jennifer Regier, Clark Wenger, and Natalie Villalobos


  • Coming Soon..


  • Tropical Ecology - Mini Project, by Lauren Taylor 2000

Entomology and Arachnology 

  • A Comparative Study of Caterpillars and Pupations in a Tropical Montane Rain Forest, by Dawn Moore 1993
  • Comparative Arthropod Survey, Shawn Dollar 1993
  • The Insect Communities of the Leaf Litter of a Primary Growth Montane Rain Forest, by Andrea Nafziger 1994
  • A comparison of Spider Diversity at the QERC, by Matthew Dykstra 1996
  • Ground Base Analysis of Lepidoptera Distribution at San Gerardo De Dota, Costa Rica, by Lúcio Mauricio Ternieden 1997
  • Butterflies of Costa Rica, by Julie Ann Birch 1998
  • Beetle Analysis, by David Hoffhines 2000


  • The Lizard Sceloporus malachiticus: an Investigation of Physical Characteristics and Behaviors of a Population in San Gerardo de Dota, Costa Rica, by Kristin Vande Griend 1995
  • A Study of the Sceloporus malachiticus of Costa Rica, by Cheri M. Pharies 1996
  • A Survey of Herpetofauna in the Savegre Reserve, by Chad A. Stallberd 2002
  • A Comparison of Preferred and Active Body Temperatures of Sceloporous malachiticus From High and Moderate Elevations in the Salamanca Cordillera of Costa Rica, by Rachel Eguren
  • Scleloporus malachiticus: A Study of Possible Navigation by Ultra Violet Light Detected Through the Parietal Eye, by Hunter Gibson


  • Lentic Algae in San Gerardo de Dota (ENG/ ES), by Nelson Aguilar 1993
  • Comparative Analysis of the Zooplankton of San Gerardo de Dota, by Dennis Siegfried 1993
  • An Inventory of Species and Interactions in the Savegre River, by Richard Sawin 1995
  • Limnology in the Rio Savegre, by Kelly Bossenbroek 1997
  • Limnology in the Savegre River, by Katherine Kingston 1998
  • Benthic Communities of a High Altitude Trout Pond, by Trina Helderman 1999
  • The Waters of the Rio Savegre Valley and the Community of San Gerardo de Dota, by Emily Hilderman 1999
  • Algae in the Savegre Valley: More Than Just Slime, by Beth Stahl 2001
  • A Chemical Analysis of the Water in the Rio Savegre, by Rachel Koss 2002
  • Water Quality in Rio Savegre, by Yuri Semenyuk 2004
  • A Comparative Study of the Inventory and Interactions of Species in the Savegre River, by Ashley A. Koskiniemi 2007


  • Diversity of Species in the Small Mammal Community in a Cloud Forest Environment at the Quetzal Education Research Complex in San Gerardo de Perez Zeldon, by Kenneth Frank Wilson 1995
  • Population Density, Interspecific Diversity and Parasitology of Small Mammals in a Mature Costa Rican Cloud Forest...A Pioneering Study, by Joe Rice 1996
  • A Methods Understanding: Camera Trap Use in the Tropics of San Gerardo de Dota, Costa Rica, by Christopher M. Tabor
  • Inferences about the Microhabitat Preferences of the Dice's Cottontail, Sylvilagus dicei, and a Study in Live Trapping, by Andrew Olsen


  • The Effects of Costa Rican Trout Farms on Coliform Levels and Antibiotic Resistance in the Lower Savegre River, by Spencer Whittenberg
  • A Survey of Coliform and Antibiotic Resistance in Bacteria Collected from the Upper Savegre Watershed in San Gerardo de Dota, by Robert Jackson


  • Density and Diversity of Mushrooms in a Primary Cloud Forest Habitat During the Dry Season, by Dawn Bakker 1996
  • Mushrooms of San Gerardo de Dota in the Rio Savegre Valley of the Talamanca Mountains, by Donna Pick 1997
  • A Survey of the Mushrooms in an Upper Montane Cloud Forest, by Anjanette Werner 1999
  • Bacteriology of Mushrooms in the Valley of San Gerardo de Dota, by Stefanie Green 2003
  • Survey of Macrofungi in San Gerardo de Dota, by Becky Schaefer 2007


  • The Spotted Wood Quail (Odontophorus guttatus), by Colin Berge 1992
  • Heterospecefic Flocking in San Gerardo de Dota, Costa Rica, by William Scott Smithson 1993
  • The Spotted Wood Quail (Odontophorus guttatus) of San Gerardo de Dota, Costa Rica, by Jason Friggens 1995
  • An Analysis of the Quetzals of San Gerardo de Dota, by John Morrow 1996
  • Sight and Sound Census of Pharomachrus mocinno and Two Other Interrelated Bird Species in the Savegre River Watershed at San Gerardo de Dota, Costa Rica, Author Unknown 1996
  • A Look at the Heterogeneous Flocking of Birds in the San Gerardo de Dota Valley, by Dave Luedeke 1996
  • Feeding Habits of the Hummingbird, by Jeff Cotner 1996
  • Bird Population Census Routes and Notes on Resplendent Quetzal Sightings, by H. Thomas Bartlett 1997
  • Observational Study of Quetzals in San Gerardo de Dota, Costa Rica, by Nathan Greenhalgh 1997
  • Nesting Survey and Correlative Study of Pharomachrus mocinno and fruiting lauraceae  in the Savegre River Watershed San Gerardo de Dota, Costa Rica, by Jeffrey Smit 1998
  • The Resplendent Quetzal: History, Habitat, and Behavior, by Jeremy Olaf Johnson 1999
  • A Study of the Nest Habitat, and Behavior for the Resplendent Quetzal, by Scott Shellenberger 2000
  • To Build a Nest: Observation fo a Colony of Long Tailed Silky-Flycatchers, by Tricia Van Dyk 2001
  • An Introductory Study of the Feeding Behaviors of the Resplendent Quetzal, by Chris Schwan 2002 
  • Interspecific Interference Competition of Six trochilidae Species in the San Gerardo de Dota Valley, by Ben Young 2002
  • Nesting Behavior Study of the Quetzal (Pharomachrus mocinno costaricensis) and the Trogonidae Family, by David Hille 2002
  • A Population Census and Behavioral Study of the Resplendent Quetzal in the Rio Savegre Valley, by Michael McNabb 2002
  • Are We Watching Too Closely?: Quetzal Behavior and Human Presence, by Heather Docter 2004
  • Preliminary Life History of the Silvery-Throated Jay (Cyanolyca argentigula), by Matthew Kuniholm 2005
  • El Nido de los Quetzales: An Observational Study of Pharomachrus micinno, by Joel Amberg 2006

Sustainable Development 

  • The Chainsaw and the White Oak, by Dr. Dwight E. Neuenschwander with Dr. Leo R. Finkenbinder 2001
  •  A Sustainable Development Solution: The Chacon Finca in the Valley of San Gerardo de Dota, by Annette Rodriguez 1994
  • A Comparative Study of Ecotourism and Sustainable Development in San Gerardo de Dota, by Dusty Watts 1994
  • Recycling in San Gerardo de Dota, by Debbie Stansell 1996
  • Agriculture Reforestation in San Gerardo de Dota, Costa Rica, by Steven Sanchez 1996
  • A Comparative Study of Sustainable Tourism Trends in San Gerardo de Dota, by Kelli Fettig 1998
  • The Fruit Production of the Savegre Valley, by Andrea Bumpass 1998
  • Ecotourism's Poster Child, by Brendan Blowers 2001
  • Sustainable Ecotourism in San Gerardo de Dota, by Marla Simmers 2002
  • Effective Interpretive Nature Trail Guide (ENG/ ES), by Kyla Buchwald 2004
  • What is the Value of a Quetzal? (ENG/ ES), by Micah Hatch 2004
  • The Sustainability of the Orchards and Trout Ponds of San Gerardo de Dota (ENG/ ES), by Blaire Knighton 2005